The oldest L. pictus fossil dates back to 200,000 years ago and was found in HaYonim Cave, Israel. The evolution of the African wild dog is poorly understood due to the scarcity of fossil finds. Some authors consider the extinct Canis subgenus Xenocyon as ancestral to both the genus Lycaon and the genus Cuon, :p149 which lived throughout Eurasia and Africa from the Early Pleistocene to the early Middle Pleistocene. Others propose that Xenocyon should be reclassified as Lycaon. The species Canis (Xenocyon) falconeri shared the African wild dog's absent first metacarpal (dewclaw), though its dentition was still relatively unspecialised. This connection was rejected by one author because C. (X. ) falconeri's missing metacarpal was a poor indication of phylogenetic closeness to the African wild dog and the dentition was too different to imply ancestry. Another ancestral candidate is the Plio-Pleistocene L. sekowei of South Africa on the basis of distinct accessory cusps on its premolars and anterior accessory cuspids on its lower premolars. These adaptions are found only in Lycaon among living canids, which shows the same adaptations to a hypercarnivorous diet. L. sekowei had not yet lost the first metacarpal absent in L. pictus and was more robust than the modern species, having 10% larger teeth.